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Welcome to theFoundation for Applied Molecular Evolution The last half century of medical research has placed chemical structures behind much of biology including human disease the human genome and the origin of life. Scientists at the Foundation have contributed broadly to these activities through innovative and polydisciplinary research in fields as diverse as chemistry informatics biology geology and astronomy. We are now taking the next steps to place biological chemistry within its larger systems context from the cell to the organism and from there to the ecosystem and the planet. Emerging from this are new tools for systems biology and personalized medicine as well as answers to some big questions Where did we come from.